August Alphonse Derbes (May 8, 1818, Marseille – January 27, 1894, Marseille) was a French professor of naturalist , zoologist and botanist at the University of Marseilles who studied reproduction of sea urchins and algae . Derbès Was the first scientist to observe the fertilization of an egg in an animal When He detailed the process of forming an envelope around the gamete During sea urchin reproduction process has now Known to be associated with Ca 2+ release.  
- Jump up^ “August Alphonse Derbès at IPNI” . Retrieved 28 July 2012 .
- Jump up^ Briggs, Elissa; Gary M. Wessel (2006). “In the beginning … Animal fertilization and sea urchin development”. Developmental Biology . 300 : 15-26. doi : 10.1016 / j.ydbio.2006.07.014 .
- Jump up^ Parrington, John; Davis, Lianne C .; Galione, Antony; Wessel, Gary (August 1, 2007). “Flipping the switch: How to sperm activates the egg at fertilization”. Developmental Dynamics . 236 (8): 2027-2038. doi : 10.1002 / dvdy.21255 . PMID 17654712 .
- Jump up^ IPNI . Derbes .